| EC |
1.2.3.1 |
| Accepted name: |
aldehyde oxidase |
| Reaction: |
an aldehyde + H2O + O2 = a carboxylate + H2O2 |
| Other name(s): |
quinoline oxidase; retinal oxidase |
| Systematic name: |
aldehyde:oxygen oxidoreductase |
| Comments: |
Contains molybdenum, [2Fe-2S] centres and FAD. The enzyme from liver exhibits a broad substrate specificity, and is involved in the metabolism of xenobiotics, including the oxidation of N-heterocycles and aldehydes and the reduction of N-oxides, nitrosamines, hydroxamic acids, azo dyes, nitropolycyclic aromatic hydrocarbons, and sulfoxides [4,6].The enzyme is also responsible for the oxidation of retinal, an activity that was initially attributed to a distinct enzyme, retinal oxidase (formerly EC 1.2.3.11) [5,7]. |
| Links to other databases: |
BRENDA, EAWAG-BBD, EXPASY, Gene, KEGG, MetaCyc, PDB, CAS registry number: 9029-07-6 |
| References: |
| 1. |
Gordon, A.H., Green, D.E. and Subrahmanyan, V. Liver aldehyde oxidase. Biochem. J. 34 (1940) 764–774. [PMID: 16747217] |
| 2. |
Knox, W.E. The quinine-oxidizing enzyme and liver aldehyde oxidase. J. Biol. Chem. 163 (1946) 699–711. [PMID: 20985642] |
| 3. |
Mahler, H.R., Mackler, B., Green, D.E. and Bock, R.M. Studies on metalloflavoproteins. III. Aldehyde oxidase: a molybdoflavoprotein. J. Biol. Chem. 210 (1954) 465–480. [PMID: 13201608] |
| 4. |
Krenitsky, T.A., Neil, S.M., Elion, G.B. and Hitchings, G.H. A comparison of the specificities of xanthine oxidase and aldehyde oxidase. Arch. Biochem. Biophys. 150 (1972) 585–599. [DOI] [PMID: 5044040] |
| 5. |
Tomita, S., Tsujita, M. and Ichikawa, Y. Retinal oxidase is identical to aldehyde oxidase. FEBS Lett. 336 (1993) 272–274. [DOI] [PMID: 8262244] |
| 6. |
Yoshihara, S. and Tatsumi, K. Purification and characterization of hepatic aldehyde oxidase in male and female mice. Arch. Biochem. Biophys. 338 (1997) 29–34. [DOI] [PMID: 9015384] |
| 7. |
Huang, D.-Y., Furukawa, A. and Ichikawa, Y. Molecular cloning of retinal oxidase/aldehyde oxidase cDNAs from rabbit and mouse livers and functional expression of recombinant mouse retinal oxidase cDNA in Escherichia coli. Arch. Biochem. Biophys. 364 (1999) 264–272. [DOI] [PMID: 10190983] |
| 8. |
Uchida, H., Kondo, D., Yamashita, A., Nagaosa, Y., Sakurai, T., Fujii, Y., Fujishiro, K., Aisaka, K. and Uwajima, T. Purification and characterization of an aldehyde oxidase from Pseudomonas sp. KY 4690. FEMS Microbiol. Lett. 229 (2003) 31–36. [DOI] [PMID: 14659539] |
|
| [EC 1.2.3.1 created 1961, modified 2002, modified 2004, modified 2012] |
| |
|
| |
|
|
EC
|
1.2.3.2
|
| Transferred entry: | xanthine oxidase. Now EC 1.17.3.2, xanthine oxidase
|
| [EC 1.2.3.2 created 1961, deleted 1984] |
| |
|
| |
|
| EC |
1.2.3.3 |
| Accepted name: |
pyruvate oxidase |
| Reaction: |
pyruvate + phosphate + O2 = acetyl phosphate + CO2 + H2O2 |
| Glossary: |
thiamine diphosphate = 3-[(4-amino-2-methylpyrimidin-5-yl)methyl]-5-(2-diphosphoethyl)-4-methyl-1,3-thiazolium |
| Other name(s): |
pyruvic oxidase; phosphate-dependent pyruvate oxidase |
| Systematic name: |
pyruvate:oxygen 2-oxidoreductase (phosphorylating) |
| Comments: |
A flavoprotein (FAD) requiring thiamine diphosphate. Two reducing equivalents are transferred from the resonant carbanion/enamine forms of 2-hydroxyethyl-thiamine-diphosphate to the adjacent flavin cofactor, yielding 2-acetyl-thiamine diphosphate (AcThDP) and reduced flavin. FADH2 is reoxidized by O2 to yield H2O2 and FAD and AcThDP is cleaved phosphorolytically to acetyl phosphate and thiamine diphosphate [2]. |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc, PDB, CAS registry number: 9001-96-1 |
| References: |
| 1. |
Williams, F.R. and Hager, L.P. Crystalline flavin pyruvate oxidase from Escherichia coli. I. Isolation and properties of the flavoprotein. Arch. Biochem. Biophys. 116 (1966) 168–176. [PMID: 5336022] |
| 2. |
Tittmann, K., Wille, G., Golbik, R., Weidner, A., Ghisla, S. and Hübner, G. Radical phosphate transfer mechanism for the thiamin diphosphate- and
FAD-dependent pyruvate oxidase from Lactobacillus plantarum. Kinetic
coupling of intercofactor electron transfer with phosphate transfer to
acetyl-thiamin diphosphate via a transient FAD
semiquinone/hydroxyethyl-ThDP radical pair. Biochemistry 44 (2005) 13291–13303. [DOI] [PMID: 16201755] |
|
| [EC 1.2.3.3 created 1961] |
| |
|
| |
|
| EC |
1.2.3.4 |
| Accepted name: |
oxalate oxidase |
| Reaction: |
oxalate + O2 + 2 H+ = 2 CO2 + H2O2 |
| Other name(s): |
aero-oxalo dehydrogenase; oxalic acid oxidase |
| Systematic name: |
oxalate:oxygen oxidoreductase |
| Comments: |
Contains Mn2+ as a cofactor. The enzyme is not a flavoprotein as had been thought [3]. |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc, PDB, CAS registry number: 9031-79-2 |
| References: |
| 1. |
Datta, P.K., Meeuse, B.J.D., Engstrom-Heg, V. and Hilal, S.H. Moss oxalic acid oxidase - a flavoprotein. Biochim. Biophys. Acta 17 (1955) 602–603. [PMID: 13250021] |
| 2. |
Kotsira, V.P. and Clonis, Y.D. Oxalate oxidase from barley roots: purification to homogeneity and study of some molecular, catalytic, and binding properties. Arch. Biochem. Biophys. 340 (1997) 239–249. [DOI] [PMID: 9143327] |
| 3. |
Requena, L. and Bornemann, S. Barley (Hordeum vulgare) oxalate oxidase is a manganese-containing enzyme. Biochem. J. 343 (1999) 185–190. [PMID: 10493928] |
|
| [EC 1.2.3.4 created 1961] |
| |
|
| |
|
| EC |
1.2.3.5 |
| Accepted name: |
glyoxylate oxidase |
| Reaction: |
glyoxylate + H2O + O2 = oxalate + H2O2 |
| Systematic name: |
glyoxylate:oxygen oxidoreductase |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc, PDB, CAS registry number: 37251-03-9 |
| References: |
| 1. |
Kasai, T., Suzuki, I. and Asai, T. [Glyoxylic oxidase system in Acetobacter.] Koso Kagaku Shimpojiumu 17 (1962) 77–81. (in Japanese) |
|
| [EC 1.2.3.5 created 1972] |
| |
|
| |
|
| EC |
1.2.3.6 |
| Accepted name: |
pyruvate oxidase (CoA-acetylating) |
| Reaction: |
pyruvate + CoA + O2 = acetyl-CoA + CO2 + H2O2 |
| Systematic name: |
pyruvate:oxygen 2-oxidoreductase (CoA-acetylating) |
| Comments: |
A flavoprotein (FAD). May be identical with EC 1.2.7.1 pyruvate synthase. |
| Links to other databases: |
BRENDA, EXPASY, KEGG, MetaCyc, CAS registry number: 62213-57-4 |
| References: |
| 1. |
Reeves, R.E., Warren, L.G., Susskind, B. and Lo, H.-S. An energy-conserving pyruvate-to-acetate pathway in Entamoeba histolytica. Pyruvate synthase and a new acetate thiokinase. J. Biol. Chem. 252 (1977) 726–731. [PMID: 13076] |
| 2. |
Takeuchi, T., Weinbach, E.C. and Diamond, L.S. Pyruvate oxidase (CoA acetylating) in Entamoeba histolytica. Biochem. Biophys. Res. Commun. 65 (1975) 591–596. [DOI] [PMID: 167776] |
|
| [EC 1.2.3.6 created 1976] |
| |
|
| |
|
| EC |
1.2.3.7 |
| Accepted name: |
indole-3-acetaldehyde oxidase |
| Reaction: |
(indol-3-yl)acetaldehyde + H2O + O2 = (indol-3-yl)acetate + H2O2 |
| Other name(s): |
indoleacetaldehyde oxidase; IAAld oxidase; AO1; indole-3-acetaldehyde:oxygen oxidoreductase |
| Systematic name: |
(indol-3-yl)acetaldehyde:oxygen oxidoreductase |
| Comments: |
A hemoprotein. This enzyme is an isoform of aldehyde oxidase (EC 1.2.3.1). It has a preference for aldehydes having an indole-ring structure as substrate [6,7]. It may play a role in plant hormone biosynthesis as its activity is higher in the auxin-overproducing mutant, super-root1, than in wild-type Arabidopsis thaliana [7]. While (indol-3-yl)acetaldehyde is the preferred substrate, it also oxidizes indole-3-carbaldehyde and acetaldehyde, but more slowly. The enzyme from maize contains FAD, iron and molybdenum [4]. |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc, CAS registry number: 66082-22-2 |
| References: |
| 1. |
Bower, P.J., Brown, H.M. and Purves, W.K. Cucumber seedling indoleacetaldehyde oxidase. Plant Physiol. 61 (1978) 107–110. [PMID: 16660220] |
| 2. |
Miyata, S., Suzuki, Y., Kamisaka, S. and Masuda, Y. Indole-3-acetaldehyde oxidase of pea-seedlings. Physiol. Plant. 51 (1981) 402–406. |
| 3. |
Rajagopal, R. Metabolism of indole-3-acetaldehyde. III. Some characteristics of the aldehyde oxidase of Avena coleoptiles. Physiol. Plant. 24 (1971) 272–281. |
| 4. |
Koshiba, T., Saito, E., Ono, N., Yamamoto, N. and Sato, M. Purification and properties of flavin- and molybdenum-containing aldehyde oxidase from coleoptiles of maize. Plant Physiol. 110 (1996) 781–789. [PMID: 12226218] |
| 5. |
Koshiba, T. and Matsuyama, H. An in vitro system of indole-3-acetic acid formation from tryptophan in maize (Zea mays) coleoptile extracts. Plant Physiol. 102 (1993) 1319–1324. [PMID: 12231908] |
| 6. |
Sekimoto, H., Seo, M., Kawakami, N., Komano, T., Desloire, S., Liotenberg, S., Marion-Poll, A., Caboche, M., Kamiya, Y. and Koshiba, T. Molecular cloning and characterization of aldehyde oxidases in Arabidopsis thaliana. Plant Cell Physiol. 39 (1998) 433–442. [PMID: 9615466] |
| 7. |
Seo, M., Akaba, S., Oritani, T., Delarue, M., Bellini, C., Caboche, M. and Koshiba, T. Higher activity of an aldehyde oxidase in the auxin-overproducing
superroot1 mutant of Arabidopsis thaliana. Plant Physiol. 116 (1998) 687–693. [PMID: 9489015] |
|
| [EC 1.2.3.7 created 1984, modified 2004, modified 2006] |
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| |
|
| EC |
1.2.3.8 |
| Accepted name: |
pyridoxal oxidase |
| Reaction: |
pyridoxal + H2O + O2 = 4-pyridoxate + (?) |
|
For diagram of pyridoxal catabolism, click here |
| Systematic name: |
pyridoxal:oxygen 4-oxidoreductase |
| Comments: |
A molybdenum protein. |
| Links to other databases: |
BRENDA, EAWAG-BBD, EXPASY, Gene, KEGG, MetaCyc, PDB, CAS registry number: 76415-81-1 |
| References: |
| 1. |
Hanly, E.W. Preliminary characterization and physical properties of pyridoxal oxidase activity from Drosophila melanogaster. Mol. Gen. Genet. 180 (1980) 455–462. |
| 2. |
Warner, C.K., Watts, D.T. and Finnerty, V. Molybdenum hydroxylases in Drosophila. I. Preliminary studies of pyridoxal oxidase. Mol. Gen. Genet. 180 (1980) 449–453. |
|
| [EC 1.2.3.8 created 1984] |
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|
| |
|
| EC |
1.2.3.9 |
| Accepted name: |
aryl-aldehyde oxidase |
| Reaction: |
an aromatic aldehyde + O2 + H2O = an aromatic carboxylate + H2O2 |
| Systematic name: |
aryl-aldehyde:oxygen oxidoreductase |
| Comments: |
Acts on benzaldehyde, vanillin and a number of other aromatic aldehydes, but not on aliphatic aldehydes or sugars. |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc, CAS registry number: 82657-93-0 |
| References: |
| 1. |
Crawford, D.L., Sutherland, J.B., Pometto, A.L., III and Miller, J.M. Production of an aromatic aldehyde oxidase by Streptomyces viridosporus. Arch. Microbiol. 131 (1982) 351–355. |
|
| [EC 1.2.3.9 created 1986, modified 2002] |
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|
|
EC
|
1.2.3.10
|
| Deleted entry: | carbon-monoxide oxidase. Activity due to EC 1.2.2.4 carbon-monoxide dehydrogenase (cytochrome b-561) |
| [EC 1.2.3.10 created 1990, deleted 2003] |
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|
|
EC
|
1.2.3.11
|
| Deleted entry: | retinal oxidase. Now included with EC 1.2.3.1, aldehyde oxidase |
| [EC 1.2.3.11 created 1990, modified 2002, deleted 2011] |
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|
|
EC
|
1.2.3.12
|
| Transferred entry: | vanillate demethylase. Now EC 1.14.13.82, vanillate monooxygenase
|
| [EC 1.2.3.12 created 2000, deleted 2003] |
| |
|
| |
|
| EC |
1.2.3.13 |
| Accepted name: |
4-hydroxyphenylpyruvate oxidase |
| Reaction: |
2 4-hydroxyphenylpyruvate + O2 = 2 4-hydroxyphenylacetate + 2 CO2 |
|
For diagram of 4-hydroxyphenylpyruvate metabolites, click here |
| Systematic name: |
4-hydroxyphenylpyruvate:oxygen oxidoreductase (decarboxylating) |
| Comments: |
Involved in tyrosine degradation pathway in Arthrobacter sp. |
| Links to other databases: |
BRENDA, EAWAG-BBD, EXPASY, KEGG, MetaCyc, CAS registry number: 78213-74-8 |
| References: |
| 1. |
Blakley, E.R. The catabolism of L-tyrosine by an Arthrobacter sp. Can. J. Microbiol. 23 (1977) 1128–1139. [PMID: 20216] |
|
| [EC 1.2.3.13 created 2000] |
| |
|
| |
|
| EC |
1.2.3.14 |
| Accepted name: |
abscisic-aldehyde oxidase |
| Reaction: |
abscisic aldehyde + H2O + O2 = abscisate + H2O2 |
|
For diagram of abscisic acid biosynthesis, click here |
| Other name(s): |
abscisic aldehyde oxidase; AAO3; AOd; AOδ |
| Systematic name: |
abscisic-aldehyde:oxygen oxidoreductase |
| Comments: |
Acts on both (+)- and (–)-abscisic aldehyde. Involved in the abscisic-acid biosynthesis pathway in plants, along with EC 1.1.1.288, (xanthoxin dehydrogenase), EC 1.13.11.51 (9-cis-epoxycarotenoid dioxygenase) and EC 1.14.14.137 [(+)-abscisic acid 8′-hydroxylase]. While abscisic aldehyde is the best substrate, the enzyme also acts with indole-3-aldehyde, 1-naphthaldehyde and benzaldehyde as substrates, but more slowly [3]. |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc, CAS registry number: 129204-36-0 |
| References: |
| 1. |
Sagi, M., Fluhr, R. and Lips, S.H. Aldehyde oxidase and xanthin dehydrogenase in a flacca tomato mutant with deficient abscisic acid and wilty phenotype. Plant Physiol. 120 (1999) 571–577. [PMID: 10364409] |
| 2. |
Seo, M., Peeters, A.J., Koiwai, H., Oritani, T., Marion-Poll, A., Zeevaart, J.A., Koornneef, M., Kamiya, Y. and Koshiba, T. The Arabidopsis aldehyde oxidase 3 (AAO3) gene product catalyzes the final step in abscisic acid biosynthesis in leaves. Proc. Natl. Acad. Sci. USA 97 (2000) 12908–12913. [DOI] [PMID: 11050171] |
| 3. |
Seo, M., Koiwai, H., Akaba, S., Komano, T., Oritani, T., Kamiya, Y. and Koshiba, T. Abscisic aldehyde oxidase in leaves of Arabidopsis thaliana. Plant J. 23 (2000) 481–488. [DOI] [PMID: 10972874] |
|
| [EC 1.2.3.14 created 2005] |
| |
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| |
|
| EC |
1.2.3.15 |
| Accepted name: |
(methyl)glyoxal oxidase |
| Reaction: |
(1) glyoxal + H2O + O2 = glyoxylate + H2O2 (2) 2-oxopropanal + H2O + O2 = pyruvate + H2O2 |
| Glossary: |
2-oxopropanal = methylglyoxal |
| Other name(s): |
glx1 (gene name); glx2 (gene name) |
| Systematic name: |
(methyl)glyoxal:oxygen oxidoreductase |
| Comments: |
The enzyme, originally characterized from the white rot fungus Phanerochaete chrysosporium, utilizes a free radical-coupled copper complex for catalysis. |
| Links to other databases: |
BRENDA, EXPASY, Gene, KEGG, MetaCyc |
| References: |
| 1. |
Kersten, P.J. and Kirk, T.K. Involvement of a new enzyme, glyoxal oxidase, in extracellular H2O2 production by Phanerochaete chrysosporium. J. Bacteriol. 169 (1987) 2195–2201. [DOI] [PMID: 3553159] |
| 2. |
Kersten, P.J. and Cullen, D. Cloning and characterization of cDNA encoding glyoxal oxidase, a H2O2-producing enzyme from the lignin-degrading basidiomycete Phanerochaete chrysosporium. Proc. Natl. Acad. Sci. USA 90 (1993) 7411–7413. [DOI] [PMID: 8346264] |
| 3. |
Kersten, P.J., Witek, C., vanden Wymelenberg, A. and Cullen, D. Phanerochaete chrysosporium glyoxal oxidase is encoded by two allelic variants: structure, genomic organization, and heterologous expression of glx1 and glx2. J. Bacteriol. 177 (1995) 6106–6110. [DOI] [PMID: 7592374] |
| 4. |
Whittaker, M.M., Kersten, P.J., Nakamura, N., Sanders-Loehr, J., Schweizer, E.S. and Whittaker, J.W. Glyoxal oxidase from Phanerochaete chrysosporium is a new radical-copper oxidase. J. Biol. Chem. 271 (1996) 681–687. [DOI] [PMID: 8557673] |
|
| [EC 1.2.3.15 created 2016] |
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